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I get the error regardless of that setting. Before I used the default port 465 but now I changed it to 25 (see http://www-01.ibm.com/support/docview.wss?uid=swg21108352). After this change I was able to receive the e-mail (!) but I still get the error and it looks like it is not using TLS. It could also be that there was a delay in the DNS update and that's why it started to work now.
Here is what I get in the log (some SMTP debug settings are enabled):
[0BA8:0006-0DA0] 25.10.2011 15:51:48.11 SMTP CIServ Listen> Connection Accepted on Port 25 for Session 13B30002
[0BA8:0008-0FC4] 25.10.2011 15:51:48.12 SMTP CITask StateMachine> Sent 106 bytes to 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:48.25 SMTP CITask StateMachine> Received 28 bytes from 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:48.25 SMTP CITask StateMachine> Sent 166 bytes to 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0DF0] State change from Greeting to Greeting
[0BA8:0008-0DF0] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0DF0] Processing in Greeting state
[0BA8:0008-0DF0] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0DF0] State change from Greeting to Connected
[0BA8:0008-0DF0] 25.10.2011 15:51:48 SMTP Server: mail-yw0-f46.google.com (209.85.213.46) connected
[0BA8:0008-0DF0] 25.10.2011 15:51:48.37 SMTP CITask StateMachine> Received 8 bytes from 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:48.37 SMTP CITask StateMachine> Sent 24 bytes to 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0FC4] Processing in Connected state
[0BA8:0008-0FC4] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0FC4] EHLO command received
[0BA8:0008-0DF0] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0DF0] Processing in Connected state
[0BA8:0008-0DF0] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0DF0] STARTTLS command received
[0BA8:0008-0FC4] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0FC4] Processing in Connected state
[0BA8:0008-0FC4] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0FC4] STARTTLS command (cont.)
[0BA8:0008-0FC4] 25.10.2011 15:51:48 ISiteAdrs Server Error: SSL Port is enabled but there are no Internet Site Documents allowing SSL for this server.
[0BA8:0008-0FC4] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0FC4] Processing in Connected state
[0BA8:0008-0FC4] 25.10.2011 15:51:48 SMTP Server [0BA8:0008-0FC4] STARTTLS command (cont.)
[0BA8:0008-0DF0] 25.10.2011 15:51:48.93 SMTP CITask StateMachine> Received 28 bytes from 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:48.93 SMTP CITask StateMachine> Sent 143 bytes to 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] Processing in Connected state
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] EHLO command received
[0BA8:0008-0DF0] 25.10.2011 15:51:49.06 SMTP CITask StateMachine> Received 29 bytes from 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] Processing in Connected state
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] MAIL command received
[0BA8:0008-0FC4] 25.10.2011 15:51:49.43 SMTP CITask StateMachine> Sent 36 bytes to 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:49.56 SMTP CITask StateMachine> Received 27 bytes from 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:49.56 SMTP CITask StateMachine> Sent 39 bytes to 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] Processing in Connected state
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] RCPT command received
[0BA8:0008-0DF0] 25.10.2011 15:51:49.70 SMTP CITask StateMachine> Received 4 bytes from 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:49.70 SMTP CITask StateMachine> Sent 53 bytes to 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] Processing in Connected state
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] DATA command received
[0BA8:0008-0DF0] 25.10.2011 15:51:49.82 SMTP CITask StateMachine> ITASK_STATE_RECV_TEXT: Received total line 32, 1 bytes from 209.85.213.46
[0BA8:0008-0FC4] 25.10.2011 15:51:49.84 SMTP CITask StateMachine> Sent 35 bytes to 209.85.213.46
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] Processing in Connected state
[0BA8:0008-0DF0] 25.10.2011 15:51:49 SMTP Server [0BA8:0008-0DF0] DATA command (cont.)
[0BA8:0008-0DF0] 25.10.2011 15:51:50 SMTP Server: Message 0046A9C6 (MessageID: <CACfGDnvKkQfSUOPCwL3Qid3KjQNpoYuwXg3VuNSWW8=nw3aQSA@mail.gmail.com>) received
[0624:0008-0844] 25.10.2011 15:51:50 Router: Message 0046A9C6 successfully moved to database xxxxx.nsf by filter action.
Feedback response number WEBB8MYH6T created by ~Alexis Minlutex on 10/25/2011